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General comparisons
Tails seems to have quite a few collision problems.
Shields do not protect you from the debris of the Iwamodokis.
Act 1
Knuckles doesn't run in at the start of Act 1. He just stands there, and there is a pile of rocks behind his starting position.
The Heat Arms has a few messed up textures.
Map
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0517 prototype
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Sonic & Knuckles final
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Act 2
Act 2's palette now changes to indicate a freeze-over when starting the Act from Act 1, albeit the lava cools more slowly. However, if the Act is accessed via the Level Select, the player dies or returns from a Bonus or Special Stage, the old palette will be loaded instead.
When you encounter Knuckles, his theme doesn't play, and he laughs after punching the boulder at you. The boulder won't cancel your Super/Hyper form.
The Act 2 boss area does not play the boss theme at the start. It plays it as soon as the boss emerges from the lava.
The Act 2 boss area does not have a Lightning Shield. The ring pattern is also different.
When you break the capsule at the end of the boss fight in Act 2, the platform on the screen floats towards the other platforms. You have to jump from platform to platform to get to Hidden Palace unlike in the final. This part is almost unchanged from the 0408 prototype except the transition to Hidden Palace does work.
Map
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0517 prototype
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Sonic & Knuckles final
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References